Checklist of the yellowjackets of the world (Vespinae)

Systematics and other topics

Checklist of the yellowjackets of the world (Vespinae)

Postby Anthony Daglio on 18 Aug 2016, 17:09

Checklist of the yellowjackets of the world (Hymenoptera; Vespinae)

Genus Pseudovespula Bischoff
Pseudovespula Bischoff, 1931 (1930), Sitz.-Ber. Ges. Naturf. Freunde Berl.: 346, subgenus of Pseudovespa Schmiedeknecht. Type species: “Ps. adulterina Buyss.”
[= Vespa norwegica var. adulterina du Buysson, 1905], by original designation.

adulterina (du Buysson)
Vespa norwegica var. adulterina du Buysson, 1905 (1904), Ann. Soc. Entomol. Fr. 73 (3): 499 (key), 600, 628, female - “Europe: Piémont; Vosges, Remiremont; Suisse, Engadine, Rosegethal, Amérique; Orégon, Corvallis” “Mongolie; Ile Sakhaline. Village de Lasevo, pès de la station de Viterbsk, Russie” (Paris, München, Washington, St.Petersburg).
Vespa norvegica saxonica natio colchica Birula, 1930, Ann. Mus. Zool. Acad. Sci. U. R. S. S. 31 (2): 314, male - [Georgia] “Adsharien (Batum-Gebiet, Alpenwiesen, s. g. Jailag, bei Sarytschair auf der Höhe von etwa 2300 m ü. d. M)” (St. Petersburg). Unavailable under Article 45.5 of the International Code of Zoological Nomenclature.
Distribution: Norway; Sweden; Finland; Denmark; Belgium; Spain, Netherlands; France; Luxembourg; Switzerland; Italy; Germany; Austria; Liechtenstein; Hungary; Croatia; Czech Republic; Poland; Slovakia; Greece; Serbia; Albania; Slovenia; Bosnia and Herzegovina; Macedonia; Romania; Moldova; Belarus; Lithuania; Latvia; Estonia; Ukraine; Russia: European Russia, Kola Peninsula, Altai, Caucasus to Siberia to Chutkotka Peninsula, Amur; Primor’ye; Sakhalin Island; Georgia; Turkey.

adulterina montivaga Yamane
Dolichovespula adulterina montivaga Yamane, 1982, Trans. Shikoku Entomol. Soc. 16: 109, 110, female, male, fig.1., - “Mt. Kariba (1500 m alt.), Hokkaido, Japan” (holotype female Kagoshima); also from Honshu. male, fig.1., - “Mt. Kariba (1500 m alt.), Hokkaido, Japan” (holotype female Kagoshima); also from Honshu.
Distribution: Japan: Hokkaido, Honshu. Russia: Kurile Islands: Kunashir.

adulterina ssp. #A
Distribution: Kazakhstan; Kyrgyzstan; Mongolia.

adulterina ssp. #B
Distribution: China: Sichuan, Shaanxi.

arctica (Rohwer)
Vespa borealis; Lewis, 1897, Trans. Am. Entomol. Soc. 24: 171 (key), 174 (female, male; “United States, New York, Mt. Washington, British Columbia; Amherst, Mass.”). Misidentification.
Vespula arctica Rohwer, 1916, Conn. State Geol. Nat. Hist. Surv. Bull. 22: 642 (key; in subgenus
Dolichovespula). New name for Vespa borealis sensu Lewis.
Distribution: Canada: Yukon Terr., N. W. T., B. C., Alta., Sask., Man., Ont., Que., N. B., Nova Scotia; P. E. I., Newfoundland; Anticosti Island; U. S. A.: AK, WA, OR, CA, NV, ID, MT, WY, UT, CO, AZ, NM, ND, MN, WI, IL, MI, IN, OH, PA, NY, VT, NH, ME, MA, RI, CT, NJ, DE, MD, VA, WV, KY, TN, NC, SC, GA.

arctica spp #A
Distribution: Canada: B.C., Alta, U.S.A.: OR, WA, CA. ID, UT, CO, WY.

arctica spp #B
Distribution: Canada: B.C., Alta, U.S.A.: OR, WA, CA. ID, UT, CO, WY.

omissa (Bischoff)
Vespa tripunctata Schenck, 1861, Jahrb. Ver. Naturk. Nassau 16: 17 (key), 25, female - [Germany] “Bamberg” (München). Junior primary homonym of Vespa 3-punctata Fabricius, 1787.
Vespula norvegica saxonica morpha ingrica Birula, 1930, Ann. Mus. Zool. Acad. Sci. U. R. S. S. 31 (2): 309, female - “Leningrader Gouvernement” (lectotype from “Leningrad, Kaipolova” in St. Petersburg). Unavailable under Article 45.5 of the International Code of Zoological Nomenclature.
Vespa omissa Bischoff, 1931, Mitt. Deutsch. Entomol. Ges. 2 (1): 6, female - [Austria] “Thüringen” (Berlin); also from Tirol.
Vespula ingrica “(Birula)” Bequaert, 1932, Entomol. Am. (n. s.) 12 (2): 89, footnote. Available under Articles 10.2, 23.3.4 and 50.3.1 of the International Code of Zoological Nomenclature.
Distribution: Norway; Sweden; Finland; Denmark; Germany; Belgium; Netherlands; Luxembourg; France; Spain; Italy; Malta; Czech Republic; Austria; Liechtenstein; Poland; Hungary; Croatia; Slovenia; Slovakia; Serbia; Bulgaria; Albania; Bosnia and Herzegovina; Macedonia; Greece; Estonia; Latvia; Lithuania; Belarus; Ukraine; Moldova; Romania; Switzerland; Russia: European Russia, Altai, Siberia; Kazakhstan; Azerbaijan; Georgia; Turkey; Iran.

Genus Dolichovespula Rohwer
Dolichovespula Rohwer, 1916, Conn. State Geol. Nat. Hist. Surv. Bull. 22: 642, subgenus of Vespula Thomson. Type species: Vespa maculata Linnaeus, 1763, by original designation.

flora Archer
Dolichovespula flora Archer, 1987, Entomol. Mon. Mag. 123: 27, female - “Burma, Adung Valley” (London); also from China.
Distribution: Myanmar; China: Gansu, Yunnan, Shaanxi, Liaoning, Sichuan.

kuami Kim & Yoon
Dolichovespula kuami Kim and Yoon, 1996, Korean J. Syst. Zool. 12 (3): 199, figs. 1, 2, female - “Korea:
Chungcheongbuk-do Province, Sokrisan” (Seoul); also from Kyeonggi-do Prov.
Distribution: Korea.

maculata (Linnaeus)
Vespa maculata Linnaeus, 1763, Amoenit. Acad. 6: 412 - [U. S. A.] “Pensylvania” (lectotype female Stockholm).
Vespa maculata americana Christ, 1791, Naturgesch. Ins.: 239 - “mitternachtlichen Amerika” (destroyed). Junior primary
homonym of Vespa americana Fabricius, 1775.
Distribution: Canada: Yukon Terr., N. W. T., B. C., Alta., Sask., Man., Ont., Que., N. B., Nova Scotia; P. E. I.,
Newfoundland; U. S. A.: AK, WA, OR, CA, NV, ID, MT, WY, UT, CO, AZ, NM, ND, SD, MN, IA, NE, KS, OK,
TX, LA, AR, MS, AL, TN, KY, MO, WI, IL, MI, IN, OH, PA, NY, VT, NH, ME, MA, RI, CT, NJ, DE, MD, VA,
WV, NC, SC, GA, FL. Introduced into New Zealand: (Dunedin, Christchurch).

media (Retzius)
Vespa crabro medius Retzius, 1783, C. Degeer Gen. Spec. Ins.: 63 (type depository unknown).
Vespa geerii Lepeletier, 1836, Hist. Nat. Ins. Hym. 1: 510, female, male - “La Suède et les environs de Paris,Versailles, dans la forêt de Saint-Germain-en-Laye” (type depository unknown).
Vespa crassa Herrich-Schäffer, 1841, Fauna Ins. Germ. 179: 33, fig. 179.4, male (destroyed).
Vespa similis Schenck, 1853, Jahrb. Ver. Naturk. Nassau 9 (1): 15 (key), 24, female, male - [Germany] “Weilburg” (Marburg?).
Vespa rufoscutellata Schenck, 1853, Jahrb. Ver. Naturk. Nassau 9 (1): 15 (key), 28, male - [Germany] “Wiesbaden” (Marburg?).
Vespa flavicincta Schenck, 1853, Jahrb. Ver. Naturk. Nassau 9 (1): 16 (key), 27, female - [Germany] “Weilburg” (Marburg?). Junior primary homonym of Vespa flavicincta Gravenhorst, 1807.
Vespa media var. lineolata Pérez, 1910, Actes Soc. Linn. Bordeaux 64: 17, female - [France] “Millau” (Paris).
Vespula media media var. lugubris Birula, 1927, Ann. Mus. Zool. Acad. Sci. U. R. S. S. 28: 80, female - “Nord-und Mittelteilen Ost-Europas, Norden” (St. Petersburg). Unavailable under Article 45.5 of the International Code of Zoological Nomenclature.
Dolichovespula media var. conjugens Paul, 1943, Stettin. Entomol. Ztg. 104: 142 - [Poland] “Stettin: Eckerberger Wald” (type depository unknown).
Dolichovespula borealis Lee, 1986, Sinozool. 10 (4): 195, 196, figs. 1(2), 2(3), female - [China] “Hengdaohezi, Heilongjiang province” (Beijing). Junior secondary homonym of Vespa borealis Kirby, 1837, and Vespa borealis Zetterstedt, 1838.
Distribution: Norway; Sweden; Finland; Denmark; Belgium; Netherlands; Luxembourg; Germany; Czech Republic; France;
Great Britain; Spain; Italy; Switzerland; Austria; Liechtenstein; Poland; Hungary; Croatia; Serbia; Slovakia;
Slovenia; Estonia; Latvia; Lithuania; Belarus; Ukraine; Moldova; Romania; Bulgaria; Albania; Bosnia and
Herzegovina; Macedonia; Greece; Turkey; Georgia; Azerbaijan; Syria; North Africa: Morocco; Russia:
European Russia south to steppe belt, Altai, Caucasus, Siberia, Yakutia, east to the Sea of Okhotsk, Kamchatka, Lake
Baikal, Amur, Primor’ye; Sakhalin Island; Kurile Islands: Iturup, Kunashir; Kazakhstan; Mongolia; China: Heilongjiang,
Liaoning, Jilin; Korea; Japan: Hokkaido. Introduced into U.K.: (England, Wales, Scotland, Isle of Man); New Zealand:
(Raglan).

Dolichovespula media sugare Ishikawa, 1969, Bull. Nat. Sci. Mus. Tokyo 12 (2): 180, figs. 1-3, female, male - “Japan
(Honshu), Tsuta, Towada, Aomori Pref.” (holotype female Shinjuku, Tokyo); also from numerous other localities on Honshu.
Distribution: Japan: Honshu.

Subgenus Boreovespula Bluthgen
Boreovespula Blüthgen, 1943, Stettin. Entomol. Ztg. 104: 149, subgenus of Dolichovespula Rohwer. Type species: “Dolichovespula norwegica (F.)” [= Vespa norwegica Fabricius, 1781], by original designation.

albida (Sladen)
Vespa marginata Kirby, 1837, Fauna Bor.-Am. 4, Ins.: 265, pl. VI fig. 2, female - [Canada] “Taken in the Route from New York, and again in Lat. 65°” (type depository unknown). Junior primary homonym of Vespa marginata Gmelin,1790.
Vespa albida Sladen, 1918, Ottawa Nat. 32: 71, male, female - [U. S. A.] “Alaska” (lectotype male Ottawa).
Distribution: Canada: Yukon Terr., N. W. T., B. C., Alta., Sask., Man., Ont., Que., N. B., Nova Scotia, P. E. I., Newfoundland; NU: Baffin Island; U. S. A.: AK, ME.

alpicola Eck
Dolichovespula alpicola “Wagner” Eck, 1984, Entomol. Abhand. Staat. Mus. Tierk. Dresden 48 (4): 40 (key), fig. 3L, female - “westliches Nordamerika”; 1987, Entomol. Abhand. Staat. Mus. Tierk. Dresden 50 (10): 191 (lectotype female from “WYO Albany County, Medicine Bow Nat. For.” in Dresden).
Distribution: Canada: Yukon Terr., B. C., Alta.; U. S. A.: AK, WA, OR, MT, WY, CO, UT, NM, AZ.

arenaria (Fabricius)
Vespa arenaria Fabricius, 1775, Syst. Entomol.: 365 - “in America arenosis” (London).
Vespa borealis Kirby, 1837, Fauna Bor.-Am. 4, Ins.: 264 - [Canada] “Lat. 65°” (type depository unknown).
Vespa diabolica de Saussure, 1854, Ét. Fam. Vesp. 2: 138, worker, queen - [U. S. A.] “L’Amérique du Nord, Philadelphie” (Torino).
Vespa fernaldi Lewis, 1897, Trans. Am. Entomol. Soc. 24: 171 (key), 173, female - [U. S. A.] “Colorado” (Philadelphia?).
Distribution: France: St. Pierre et Miquelon; Canada: Yukon Terr., N. W. T., B. C., Alta., Sask., Man., Ont., Que., N. B., Nova Scotia; P. E. I., Newfoundland; Anticosti Island; U. S. A.: AK, WA, OR, CA, NV, ID, MT, WY, UT, CO, AZ, NM, NE, ND, SD, MN, IA, WI, IL, MI, IN, OH, PA, NY, VT, NH, ME, MA, RI, CT, NJ, DE, MD, VA, WV, KY, TN, NC, SC, GA. Introduced into Bermuda.

lama (du Buysson)
Vespa lama du Buysson, 1903, Bull. Soc. Entomol. Fr. 1903: 175, female - [India] “Sikkim” (Paris).
Dolichovespula nyalamensis Lee, 1986, Sinozool. 10 (4): 195, 197, figs. 1(4), 2(2), female - [China] “Nelam, Xizang (Tibet), 3300 m” (Beijing).
Distribution: India: Uttarakhand, Sikkim; Nepal; China: Tibet.

loekenae Eck
Dolichovespula loekenae Eck, 1980, Reichenbachia 18 (30): 213, female, male - [Norway] “Espeland, HOy: Alversund” (Bergen); also from Sweden; Finland.
Distribution: Norway; Sweden; Finland; Russia: northwestern region, Ob’ River to east of Lake Baikal, Altai, Ural, Siberia, Amur, Primor’ye, Kamchatka; Kazakhstan; Kyrgyzstan; Mongolia; Korea.

norvegicoides (Sladen)
Vespa norvegicoides Sladen, 1918, Ottawa Nat. 32: 71, male, female - [Canada] “N.S. to B.C.” (lectotype female from “Amherst, Nova Scotia” in Ottawa).
Distribution: France: St. Pierre et Miquelon; Canada: Yukon Terr., N. W. T., B. C., Alta., Sask., Man., Ont., Que., N. B., Nova Scotia; P. E. I., Newfoundland; U. S. A.: AK, WA, OR, CA, ID, MT, WY, ND, MN, WI, MI, NY, VT, NH, ME, MA, RI, CT, NJ, PA, MD, VA, WV, KY, TN, NC, SC, GA.

norwegica (Fabricius)
Vespa norwegica Fabricius, 1781, Spec. Ins. 1: 460 - “in Norwegia” (Kobenhavn).
Vespa norvegica Olivier, 1792, Encycl. Méthod. Ins. 6: 674; Smith, 1858, Cat. Br. Fossor. Hym., Form. Vesp.: 220. Unjustified emendation of Vespa norwegica Fabricius.
Vespa britannica Leach, 1814, Zool. Misc. 1: 112, pl. 50, female, male - “in Britannia; in Anglia rarius, in Caledonia vulgatissime” (London).
Vespa borealis Zetterstedt, 1838, Ins. Lappon. Descr.: 454, female - “Lapponia” (? Lund). Junior primary homonym of Vespa borealis Kirby, 1837.
Vespa saxonica var. arctica Friese, 1919, Mém. Acad. Sci. Russ. (8) 28 (15): 1, female, male - [Russia] “Chanema-Fluss, Sobj-Tal, Longot-johan-Tal, Berezov” (St. Petersburg). Junior secondary homonym of Dolichovespula arctica (Rohwer, 1916).
Dolichovespula norwegica var. Zetterstedti Blüthgen, 1937, Mitt. Entomol. Ges. Halle 15: 37. Replacment name for Vespa saxonica var. arctica Friese.
Distribution: Iceland; Norway; Sweden; Finland; Denmark; Netherlands; Luxembourg; France; U. K.: England, Wales including Bardsey Island, Isle of Man, Scotland including Orkney and Shetland (Fair Isle); Ireland; Spain; Belgium; Germany; Switzerland; Liechtenstein; Italy, including Sardinia; Austria; Hungary; Croatia; Serbia; Romania; Slovakia; Poland; Czech Republic; Bulgaria; Slovenia; Albania; Bosnia and Herzegovina; Macedonia; Greece; Ukraine; Estonia; Belarus; Moldova; Latvia; Lithuania; Russia: European Russia, Altai, Caucasus, Siberia east to Chukotka and Kamchatka; Georgia; Turkey; Kazakhstan; Kyrgyzstan; Mongolia.

norwegica peruana (Saussure)
Vespa peruana de Saussure, 1867, Reise Novara., Zool. 2 (1), Hym.: 18, female – “Peru altior; Quito” [error] (Genève).
Distribution: Russia: Altai; Siberia east to Chukotka and Kamchatka; Sakhalin Island; Kazakhstan; Kyrgyzstan; China: Xinjiang; Mongolia.

pacifica (Birula)
Vespula norvegica pacifica Birula, 1930, Ann. Mus. Zool. Acad. Sci. U. S. S. R. 31: 295,317, female, male - [Russia] “Im Küstengebiete des Pazifischen Ozeans und auf den naheliegeneden Inseln” (lectotype female from “Insel Sachalin” in St. Petersburg).
Distribution: Russia: Kurile Islands: Paramushir, Shikotan, Kunashir, Iturup; Sakhalin Island; Japan: Hokkaido, Honshu, Shikoku.

panda Archer
Dolichovesupula [!] panda Archer, 1981 (1980), Entomon 5 (4): 341, fig. 1, female [male = worker] - “CHINA, Szechuen, 14.4 km south-west Tarsienlu, 2600-4000 m” (Washington); also from other localities in Sichuan.
Distribution: China: Sichuan, Ningxia, Tibet.

saxonica (Fabricius)
Vespa saxonica Fabricius, 1793, Entomol. Syst. 2: 256 - “in Germania” (type depository unknown).
Vespa bavarica von Schrank, 1802, Fauna Boica 2 (2): 350, female - [Germany] “Waldungen” (type depository unknown).
Vespa tridens Schenck, 1853, Jahrb. Ver. Naturk. Nassau 9 (1): 18 (key), 38, female, male - [Germany] “in der Gegend von Wiesbaden” (Marburg?).
Vespula norvegica saxonica morpha monticola Birula, 1930, Ann. Mus. Zool. Acad. Sci.
U. R. S. S. 31 (2): 313 - [Russia] “Gebirgen des Kaukasus” (St. Petersburg). Unavailable under Article 45.5 of the International Code of Zoological Nomenclature.
Distribution: Norway; Sweden; Finland; Denmark; Germany; Netherlands; Luxembourg; Belgium; France; Spain; Switzerland; Austria; Liechtenstein; Slovenia; Bulgaria; Italy; Greece; Poland; Hungary; Czech Republic; Slovakia; Croatia; Serbia; Estonia; Romania; Bosnia and Herzegovina; Macedonia; Albania; Latvia; Lithuania; Belarus; Ukraine; Moldova; Russia: European Russia; Paramushir; Azerbaijan; Georgia; Turkey. Introduced into U.K.: England, Wales.

saxonica nipponica Yamane
Dolichovespula saxonica nipponica Yamane, 1975, Kontyû 43 (3): 349, female - [Japan] “Hokkaido. Otoineppu” (Sapporo); also from elsewhere in Hokkaido, and Honshu.
Distribution: Japan: Hokkaido, Honshu; Russia: Kurile Islands: Kunishir.

saxonica kamtchatkensis Eck
Dolichovespula saxonica kamtschatkensis Eck, 1983, Entomol. Abhand. Staat. Mus. Tierk. Dresden 46 (8): 171, female, male - [Russia] “Petropawlowsk, Flußtal Poganka, Kamtschatka” (holotype female St. Petersburg); also from elsewhere in Kamchatka.
Distribution: Russia: Kamchatka, Komandorski Island; Kurile Islands: Paramushir.

saxonica nigrescens Eck
Dolichovespula saxonica nigrescens Eck, 1983, Entomol. Abhand. Staat. Mus. Tierk. Dresden 46 (8): 172, female - [Russia] “Primorje, Umgeb. Anisimowka” (Dresden); also from elsewhere in Russia; Mongolia; Korea.
Distribution: Kazakhstan; Mongolia; China: Xinjiang, Gansu, Heilongjiang, Ningxia, Qinghai, Sichuan, Jilin, Liaoning, Shaanxi; Korea; Russia: Altai, Ural, Siberia east to Upper Kolyma, Amur, Primor'ye, Kurile Islands: Kunashir; Sakhalin Island.

stigma Lee
Dolichovespula stigma Lee, 1986, Sinozool. 10 (4): 195, 198, figs. 1(3), 2(1), female - [China] “Mt. Emei, Sichuan Province, 1800-2100 m” (Beijing); also from 1800-1900 m.
Dolichovespula baileyi Archer, 1987, Entomol. Mon. Mag. 123: 30, female - “China, Szechwan, Lin Yang, 2500 m” (Edinburgh); also from another locality in Sichuan.
Dolichovespula sinensis Archer, 1987, Entomol. Mon. Mag. 123: 29, female - “China, Szechwan, Suifu” (Washington); also from Xinjiang.
Distribution: China: Sichuan, Xinjiang, Shaanxi.

xanthicincta Archer
Dolichovespula pacifica xanthicincta Archer, 1981 (1980), Entomon 5 (4): 341, 343, female [male = female] - “China-Tibet border, near Tang Gu, 4300 m” (Washington); also from Tibet; Myanmar.
Distribution: Bhutan; China: Sichuan, Tibet; Myanmar.

Subgenus Metavespula Bluthgen
Metavespula Blüthgen, 1943, Stettin. Entomol. Ztg. 104: 149, subgenus of Dolichovespula Rohwer.Type species: “Dolichovespula silvestris (Scop.)” [!] [= Vespa sylvestris, Scopoli, 1763], by original designation.

intermedia (Birula)
Vespula sylvestris var. intermedia Birula, 1930, Ann. Mus. Zool. Acad. Sc. U. S. S. R. 31: 330, 331, female, male - [Uzbekistan] “Zeravschan-Tal und der östlichen Buchara” (lectotype female from “Ost- Buchara, im Gebirgsrücken Peter des Grossen, Engpass Kamtschirák” in St. Petersburg).
Dolichovespula asiatica Archer, 1981, Entomol. Scand. 12: 187, 189. Unnecessary replacement name for Vespula sylvestris var. intermedia Birula, incorrectly stated to be a junior primary homonym of Vespa rufa var. intermedia du Buysson, 1905.
Distribution: Afghanistan; Turkmenistan; Uzbekistan; Tajikistan; Kazakhstan; Kyrgyzstan; Russia: Altai; Pakistan; India: Himachal Pradesh, Jammu and Kashmir; Mongolia; China: Xinjiang, Gansu.

sylvestris (Scopoli)
Vespa sylvestris Scopoli, 1763, Entomol. Carniol.: 309 - [Slovenia] “Carnioliae” (destroyed?).
Vespa parietum Harris, 1776, Exposit. Eng. Ins.: 128, pl. XXXVII fig. 4, male - “English” (destroyed). Junior primary homonym of Vespa parietum Linnaeus, 1758.
Vespa holsatica Fabricius, 1793, Entomol. Syst. 2: 257 - [Germany] “Kiliae” (Kobenhavn).
Vespa frontalis Latreille, 1802, Ann. Mus. Hist. Nat. 1: 290, pl. XXI fig. 7 - [France] “Chartreaux de Paris” (type depository unknown).
Vespa campanaria Fowler, 1833, Ann. Mag. Nat. Hist. 6: 536, 538, fig. 69 (type depository unknown).
Vespa anglica “Leach” Smith, 1843, Zoologist 1: 168, as a synonym of Vespa holsatica Fabricius, 1793. Available under Article 11.6.1 of the International Code of Zoological Nomenclature.
Vespa pilosella Costa, 1858, Ric. Entomol. Monti Partenii: 20, 28, female, male - [Italy] “Monti Partenii” (type depository unknown). Junior primary homonym of Vespa pilosella Gravenhorst, 1807.
Dolichovespula xinjiangensis Lee, 1986, Sinozool. 10 (4): 195, 199, figs. 1(1), 2(4), female, male - [China] “Xinyuan, Xinjiang Uygur Aut. Reg., 850-1200 m” (holotype female Beijing); also from Altay.
Distribution: Norway; Sweden; Finland; Denmark; Netherlands; Luxembourg; Germany including Mellum Island; Belgium; France; U. K.: England: including Isle of Man, Isles of Scilly, and the Channel Islands, Wales, Scotland including Outer Hebrides; Ireland; Portugal; Spain; Italy; Malta; Czech Republic; Switzerland; Austria; Liechtenstein; Slovenia; Slovakia; Greece; Poland; Hungary; Croatia; Serbia; Albania; Bosnia and Herzegovina; Macedonia; Bulgaria; Estonia; Latvia; Lithuania; Belarus; Ukraine; Moldova; Romania; Russia: European Russia, Altai, Caucasus, Siberia to Transbaikalia, Primor’ye; Georgia; Azerbaijan; Syria; North Africa: Morocco; Turkey; Armenia; Iran; Afghanistan; Uzbekistan; Turkmenistan; Tajikistan; Pakistan; India: Kashmir; Kazakhstan; Kyrgyzstan; Mongolia; China: Xinjiang, Gansu, Fujian, Shanghai, Heilongjiang.

sylvestris sumptuosa (Buysson)
Vespa silvestris [!] var. sumptuosa du Buysson, 1905 (1904), Ann. Soc. Entomol. Fr. 73 (3): 493 (key), 603, female - “Perse centrale” (Wien).
Distribution: Turkey; Iran; Armenia; Afghanistan; Uzbekistan; Turkmenistan; Tajikistan.

Genus Vespula Thompson
Vespula Thomson, 1869, Opusc. Entomol. 1: 79, subgenus of Vespa (8 species). Type species: Vespa austriaca Panzer, 1799, by subsequent designation of Ashmead, 1902, Can. Entomol. 34: 164.
Pseudovespa Schmiedeknecht, 1881, Entomol. Nachr. 7: 314, subgenus of Vespa Linnaeus. Type species: Vespa austriaca Panzer, 1799, by monotypy.

austriaca (Panzer)
Vespa austriaca Panzer, 1799, Faunae Ins. German. 63: 2, fig. 2, male - “Viennae Austr.” (destroyed).
Vespa biloba Schilling, 1850, Jahresber. Schles. Ges. Vaterländ. Kultur 28: 77 - [Poland] “Schlesien” (type depository unknown).
Distribution: Norway; Sweden; Finland; Denmark; Germany; Belgium; Netherlands; Luxembourg; France; Spain; Italy; Switzerland; Liechtenstein; Austria; Czech Republic; Hungary; Croatia; Serbia; Bulgaria; Greece; Albania; Bosnia and Hezegovina; Macedonia; Poland; Estonia; Latvia; Lithuania; Belarus; Ukraine; Moldova; Romania; Slovenia; Slovakia; Turkey; Georgia; Russia: European Russia, north to Arkhangel’sk, south to Moscow, Altai, Caucasus, Siberia, Kamchatka, Ussuriland; Amur, Primore’ye; Sakhalin Island; Kazakhstan; Kyrgyzstan; Mongolia; Korea.

austriaca borealis (Smith)
Vespa borealis Smith, 1843, Zoologist 1: 170, fig., female - “Yorkshire, the north of Scotland” (London). Junior primary homonym of Vespa borealis Kirby, 1837, and of Vespa borealis Zetterstedt, 1840.
Vespa arborea Smith, 1849, Zoologist 7, App.: lx. Replacement name for Vespa borealis Smith.
Distribution: U. K.: England, Wales, Scotland; Ireland.


austriaca spp #A
Vespula shinanoensis: Takamizawa, 2005, The Japanese Social Wasps and Bees: 147, 148 (in subgenus Vespula). Nomen nudum, described without mentioning repository of the holotype (Articles 13 and 16.4 of the International Code of Zoological Nomenclature).
Distribution: Japan: Hokkaido, Honshu; Russia: Sakhalin Island; Kurile Islands: Iturup, Kunashir.

austriaca spp #B
Distribution: Pakistan; India: Jammu and Kashmir; China: Hebei, Xinjiang, Shanghai, Liaoning, Shaanxi.

infernalis (de Saussure)
Vespa infernalis de Saussure, 1854, Ét. Fam. Vesp. 2: 139, worker [female] - [U. S. A.] “L’Amérique du Nord. Philadelphie” (Torino).
Vespa tripunctata Packard, 1870, Trans. Chicago Acad. Sci. 2: 26, pl. II fig. 11, female - [U. S. A.] “Kutleet” (type depository unknown). Junior primary homonym of Vespa tripunctata Fabricius, 1787, and of Vespa tripunctata Schenck, 1861.
Distribution: Canada: Yukon Terr., N. W. T., B. C., Alta., Sask., Man., Ont., Que., N. B., Nova Scotia, P. E. I., Newfoundland; U. S. A.: AK, WA, OR, CA, ID, MT, CO, UT, AZ, NM, MN, ND, NM, WI, MI, NY, NJ, VT, NH, ME, MA, RI, CT.

Genus Allovespula Bluthgen
Allovespula Blüthgen, 1943, Stettin. Entomol. Ztg. 104: 149, subgenus of Paravespula Blüthgen. Type species: “Paravespula rufa (Linné)” [= Vespa rufa Linnaeus, 1758], by monotypy.

acadica (Sladen)
Vespa rufa var. americana du Buysson, 1905 (1904), Ann. Soc. Entomol. Fr. 63: 499 (key), 592, male - “Canada, Lévis près Québec” (type depository unknown). Junior primary homonym of Vespa americana Fabricius, 1775, and of Vespa maculata americana Christ, 1791.
Vespa acadica Sladen, 1918, Ottawa Nat. 32: 72, male, female - [Canada] “Ottawa” (lectotype female Ottawa) also from Nova Scotia, N.B., B.C.
Vespula rufa var. sladeni Bequaert, 1932, Entomol. Am. (n. s.) 12 (2): 80 (key), 98, 102, female, male - [U. S. A.] “Washington State” (holotype female Cambridge); also from OR, CA, WY, UT, CO; Canada: BC.
Distribution: Canada: Yukon Terr., N. W. T., B. C., Alta., Sask., Man., Ont., Que., N. B., Nova Scotia; P. E. I., Newfoundland; U. S. A.: AK, WA, OR, CA, NV, ID, MT, WY, UT, CO, AZ, NM, NE, SD, ND, MN, WI, MI, PA, NY, VT, NH, ME, MA, RI, CT, NJ, MD, VA, WV, NC.

atropilosa (Sladen)
Vespa atropilosa Sladen, 1918, Ottawa Nat. 22: 72, female - [Canada] “Lethbridge, Alta.” (lectotype Ottawa); also from B.C.
Distribution: Canada: B. C., Alta., Sask.; U. S. A.: WA, OR, CA, NV, ID, MT, ND, NE, WY, UT, CO, AZ, NM; Mexico: Baja California Norte.

consobrina (de Saussure)
Vespa consobrina de Saussure, 1854, Ét. Fam. Vesp. 2: 141, worker - [Canada] “L’île de Terre-Neuve” (Paris).
Vespa arenaria de Saussure, 1854, Ét. Fam. Vesp. 2: 134, worker - [North America]. Junior primary homonym of Vespa arenaria Fabricius 1775.
Vespa scelesta McFarland, 1888, Trans. Am. Entomol. Soc. 15: 297 (key), 298, female, male - [U. S. A.] “Montana” (lectotype female Philadelphia); also from PA, VA, NH, CO, ME, WA, MA.
Vespa sulcata; Howard, 1901, Insect Book: pl. VI fig. 18. Lapsus for Vespa scelesta McFarland.
Distribution: France: St. Pierre et Miquelon; Canada: Yukon Terr., N. W. T., B. C., Alta., Sask., Man., Ont., Que., N. B., Nova Scotia, P. E. I., Newfoundland; U. S. A.: AK, WA, OR, CA, ID, MT, WY, UT, CO, ND, MN, WI, IL, MI, IN, OH, PA, NY, VT, NH, ME, MA, RI, CT, NJ, DE, MD, VA, WV, KY, TN, NC, SC, GA.

intermedia (du Buysson)
Vespa rufa var. intermedia du Buysson, 1905 (1904), Ann. Soc. Entomol. Fr. 73: 499 (key), 591, female, male - “Mandchourie, région du lac Hanka; Amérique du Nord: Hudson’s Bay” (Paris?, London).
Distribution: Canada: Yukon Terr., N. W. T., B. C., Alta., Sask., Man., Ont., Que., N. B., Nova Scotia, P. E. I., Newfoundland; U. S. A.: AK, ME, VT, NY.

kingdonwardi Archer
Vespula kingdonwardi Archer, 1981, Kontyû 49 (1): 54, 57 (key), 60, fig. 4C-D, female, male - [China] “Tibet, 28.25 N 97.55 E” (holotype female London); also from other localities in Tibet; Myanmar.
Vespula hirsuta Lee, 1986, Sinozool. 10 (4): 201, 204, 206, fig. 4, female - [China] “Markham County, Xizang (Tibet) Aut. Reg., 3300 m” (Beijing).
Distribution: China: Tibet, Xizang; Nepal; Myanmar.

nursei Archer
Vespula nursei Archer, 1981, Kontyû 49 (1): 54, 57, fig. 4A-B, female - “India, Kashimir, 8-9,000 ft” (London); also from other localities in Kashmir, Punjab, Himachal Pradesh; China; Philippines.
Distribution: India: Kashmir, Himachal Pradesh, Punjab; Pakistan; China: Hebei, Fujian. Introduced into Philippines: (Luzon).

rufa (Linnaeus)
Vespa rufa Linnaeus, 1758, Syst. Nat., Ed. 10, 1: 572 - “in Europa” (lectotype female, Linnaean Society, London).
Distribution: Iceland; Norway; Sweden; Finland; Denmark; Germany; Netherlands; Belgium; Luxembourg; Liechtenstein; Slovenia; Slovakia; France; Spain; Italy; Switzerland; Austria; Czech Republic; Poland; Hungary; Croatia; Serbia; Bulgaria; Albania; Bosnia and Herzegovina; Macedonia; Greece; Estonia; Latvia; Lithuania; Belarus; Moldova; Romania; Ukraine; Georgia; Azerbaijan; Turkey; Russia: European Russia north to Kola Peninsula, Arkhangel’sk and river Pechora, south to Volgograd and Orenberg.

rufa grahami Archer
Vespula rufa grahami Archer, 1981, Kontyû 50 (1): 54, 62, female, male - “China, Szechuen, 30 miles north of Tatsienku” (Washington); also from elsewhere in Sichuan.
Vespula obscura Lee, 1986, Sinozool. 10 (4): 201, 205, 206, fig. 5, female - [China] “Zogang County, Xizang (Tibet) Aut. Reg., 3800 m” (Beijing); also from another locality in Tibet.
Vespula yulongensis Dong and Wang, 2002, in Dong et al., J. Southwest Agric. Univ. 24 (5): 396, figs. 1- 5, female - “Yunnan China” (Yunnan).
Distribution: China: Yunnan, Sichuan, Tibet.

rufa spp#A
Distribution: Iceland; Norway; Sweden; Finland; Denmark; France; Russia: European Russia north to Kola Peninsula, Arkhangel’sk and river Pechora, south to Volgograd and Orenberg; Altai, Caucasus, Siberia, Kamchatka, Ussuriland, Amur, Primor’ye; Kazakhstan.

rufa ssp#B
Distribution: U. K.: England including Isle of Man and the Channel Islands, Wales including Bardsey Island, Scotland including the Outer Hebrides; Ireland.

schrenkii (Radoszkowski)
Vespa schrenckii Radoszkowski, 1861, Horae Soc. Entomol. Ross. 1: 84, figs. 4-5 - [Russia] “Amur” (type depository unknown).
Vespa sibirica André, 1884, Spec. Hym. Europe Algér. 2: 599, female, male - [Russia] “Sibérie” (Paris).
? Vespula yichunensis Lee, 1986, Sinozool. 10 (4): 202, 203, 206, fig. 3, female - [China] “Yichun City, Heilongjiang Province” (Beijing).
Distribution: Russia: Altai, Caucasus, Siberia, Kamchatka, Ussuriland, Amur, Primor’ye; Kazakhstan; Kyrgyzstan; Mongolia; China: Manchuria, Xinjiang, Yunnan, Sichuan, Heilongjiang, Hebei, Liaoning, Shaanxi, Tibet; Korea; Ulleungdo Island.

schrenkii spp #A
Distribution: Japan: Shikoku, Hokkaido, Honshu; Sakhalin Island; Kurile Islands: Urup, Iturup, Kunashir.

schrenkii spp # B
Distribution: Taiwan.

schrenkii ssp #C
Distribution: Russia: Siberia, Kamchatka, Ussuriland, Amur; Kazakhstan; Kyrgyzstan; Afghanistan; Tajikistan; Uzbekistan; Mongolia; China: Manchuria.

vidua (de Saussure)
Vespa vidua de Saussure, 1854, Ét. Fam. Vesp. 2: 136, worker - [U. S. A.] “La Caroline” (Paris).
Distribution: Canada: Man., Ont., Que, N. B., Nova Scotia; U. S. A.: ND, SD, MN, IA, WI, IL, MI, IN, OH, PA, NY, VT, NH, ME, MA, RI, CT, NJ, DE, MD, VA, WV, KY, TN, NC, SC, GA.

(New Genus: ???) Lineovespula Daglio
Type species: Vespula squamosa Drury 1773, by original designation.

squamosa (Drury)
Vespa squamosus Drury, 1773, Illustr. Nat. Hist. 2: Index to vol. I [pl. XLIII fig. 7 in Vol. 1, female - “New York”] (destroyed?).
Vespa lineata Fabricius, 1775, Syst. Entomol.: 365 - “in America” (type depository unknown). Sphex conchacea Christ, 1791, Naturgesch. Ins.: 259, pl. 25 fig. 5 - [U. S. A.] “Neuiork” (destroyed). Vespa cuneata Fabricius, 1804, Syst. Piez.: 258 - [U. S. A.] “in Carolina” (Kobenhavn).
Vespa cruciata Lepeletier, 1836, Hist. Nat. Ins. Hym. 1: 514. Unjustified emendation of Vespa cuneata Fabricius.
Vespa bistriata McFarland, 1888, Trans. Am. Entomol. Soc. 15: 297 (key), 298, female - “North America” (Philadelphia) Junior primary homonym of Vespa bistriata Fabricius, 1804.
Vespa macfarlandi Lewis, 1897, Trans. Am. Entomol. Soc. 24: 172 (key), 180. Replacement name for Vespa bistriata McFarland.
Distribution: U. S. A.: TX, LA, AR, MS, AL, OK, NE, KS, MO, IA, WI, IL, IN, MI, OH, KY, TN, WV, VA, PA, NY, NJ, DE, MD, NC, SC, GA, AL, FL; Mexico: Guerrero, Oaxaca,Tamaulipas, Chihuahua. Introduced into: Canada: Ontario; Dutch Carribean: (Curacao; St. Eustatius).

squamosa michoacana Bequaert
Vespula squamosa var. (or subsp.) michoacana Bequaert, 1941, Entomol. News 52: 249, female - “Tancitaro, 6600 ft., State of Michoacan, Mexico” (Cambridge).
Distribution: Mexico: Nuevo León,Vera Cruz, Michoacan, Chiapas, Hidalgo, Zacatecas, Puebla, México, San Luis Potosí; Guatemala; Honduras.

sulphurea (de Saussure)
Vespa sulphurea de Saussure, 1854, Ét. Fam. Vesp. 2: 137, queen - [U. S. A.] “La Californie” (London).
Distribution: U. S. A.: CA, OR, NV, AZ; Mexico: Baja California Norte.

Genus Paravespula Bluthgen
Paravespula Blüthgen, 1938 (1937), Konowia 16: 271, subgenus of Dolichovespula Rohwer. Type species: Vespa vulgaris Linnaeus, 1758, by original designation.

akrei Landolt
Vespula akrei Landolt, 2010, in Landolt et al., Zootaxa 2629: 61, 62, figs. 1A-D, female, male - “Guatemala, Zacapa, 12 km north of San Lorenzo, N15°06.86’ W89°40.82’, 2200 m. elev.” (holotype female Guatemala City); also from four other localities.
Distribution: Guatemala.

alascensis (Packard)
Vespa alascensis Packard, 1870, Trans. Chicago Acad. Sci. 2: 27, pl. II fig. 10, female - [U. S. A.] “Lower Yukon” (type depository unknown).
Vespa westwoodii Shipp, 1893, Psyche 6: 450 - “N. Amer. Bor.” (holotype female Oxford). - Dalla Torre, 1904, Gen. Ins. 19: 66 (Westwoodi [!]).
Distribution: France: St. Pierre et Miquelon; Canada: Yukon Terr., N. W. T., B. C., Alta., Sask., Man., Ont., Que., N. B., Nova Scotia, P. E. I., Newfoundland; U. S. A.: AK, WA, OR, CA, NV, ID, MT, WY, CO, UT, AZ, NM, KS, SD, ND, MN, WI, NE, IA, IL, IN, MI, OH, PA, NY, VT, NH, ME, MA, RI, CT, NJ, DE, MD, VA, WV, KY, TN, NC, SC, GA; Mexico: Durango, Michoacan, México, Oaxaca. Introduced into Hawaii: (Maui) (U.S.A.); Cuba: (Mayari Arriba).

flaviceps (Smith)
Vespa flaviceps Smith, 1870, in Horne and Smith, Trans. Zool. Soc. Lond. 7 (3): 174, 191, pl. XXI figs. 10-11, female - “Binsur, Kumaon, North-west Provinces of India” (London).
Vespa vulgaris var. flavior Stolfa, 1934, Bull. Soc. Venez. Stor. Nat. 1: 49, female - “India: Bagarkote 8000 piedi. Kumaon W. Himalaia, Dal, sopra Dharmsala, 5500 piedi Punjab, Pindi Point, Murree 7242 piedi. Punjab, Rive del fiume Deo Gad e Foresta S. O. di Pinath, 7 miglia da Kausani Distr. Almora” (type depository unknown).
Vespula japonica pionganensis Giordani Soika, 1976, Ann. Hist.-Nat. Mus. Natl. Hung. 68: 287, 290, female (in subgenus Vespula) - [Korea] “Za-mo san, 60 km NE from Pyongyan” (Budapest); also from other localities in Korea.
Vespula gracilia Lee, 1986, Sinozool. 10 (4): 201, 203, 206, fig. 2, female - “Zhoushan, Zhejiang Province” (Beijing); also from Fujian.
Distribution: Pakistan; India: Jammu and Kashmir, Punjab, Meghalaya, Manipur, Uttarakhand, Uttar Pradesh, Himachal Pradesh, Assam, Sikkim; Nepal; Myanmar; Thailand; Laos; Vietnam; China: Tibet, Sichuan, Hunan, Hebei, Hubei, Guizhou, Liaoning, Henan, Yunnan, Shaanxi, Heilongjiang, Jilin, Jiangsu, Beijing, Fujian, Zhejiang, Guangdong, Hong Kong; Russia: Amur, Primor’ye; Korea, Including Ulleungdo Island.

flaviceps lewisii
Vespa japonica de Saussure, 1858, Rev. Mag. Zool. (2) 10: 261, female - “Japon.” (Genève). Junior primary homonym of Vespa japonica Radoszkowski, 1857.
“Vespa lewisii, Sauss. Guêpes Soc. (MS.)” Smith, 1873, Trans. Entomol. Soc. Lond. 1873 (2): 198. Nomen nudum.
Vespa “lewisii, Saus. MS.” Cameron, 1903, Entomologist 36: 280, as a synonym of Vespa japonica de Saussure, 1858. Available under Article 11.6.1 of the International Code of Zoological Nomenclature.
Vespa saussurei Schulz, 1906, Spolia Hym.: 231. Replacement name for Vespa japonica de Saussure.
Distribution: Japan: Kyushu, Shikoku, Honshu, Hokkaido, Ryukyu Islands: Amami Island. Russia: Sakhalin Island.

flaviceps karenkona Sonan
Vespa karenkona Sonan, 1929, Trans. Nat. Hist. Soc. Formosa. 19 (101): 137 (key), 148, female - [Taiwan] “Karenkô” (Taichung).
Vespa 4-maculata Sonan, 1929, Trans. Nat. Hist. Soc. Formosa. 19 (101): 137 (key), 148, female - [Taiwan] “Arisan” (coll. M. Maki).
Distribution: Taiwan.

flavopilosa Jacobson
Vespula flavopilosa Jacobson, 1978, in Jacobson et al., Ann. Entomol. Soc. Am. 71: 299, 301 (key), 303, figs. 1a, 2c, ra, 5c-d, j, 8-9, 17, 22-29, table 1 (electrophoretic data), female, male - [U. S. A.] “Ithaca, Tompkins Co., NY” (holotype female Washington); also from elsewhere in eastern U.S. and Canada.
Distribution: Canada: Man., Ont., Nova Scotia; NB; U. S. A.: ND, SD, NE, MN, IA, MO, WI, IL, IN, MI, OH, PA, NY, VT, NH, ME, MA, CT, KY, NJ, DE, MD, DC, VA, WV, KY, TN, NC, SC, GA.

germanica (Fabricius)
Vespa germanica Fabricius, 1793, Entomol. Syst. 2: 256 - [Germany] “Kiliae” (type depository unknown).
Distribution: Norway; Sweden; Finland; Denmark; Spain including Canary Islands; Portugal including Azores; Malta; Netherlands; Belgium; Luxembourg; Germany; U. K.: England including Channel Islands and Isles of Scilly, Wales, Scotland including Shetland; Ireland; Scotland; France; Switzerland; Austria; Liechtenstein; Czech Republic; Hungary; Slovenia; Slovakia; Croatia; Serbia; Bulgaria; Albania; Bosnia and Herzegovina; Macedonia; Italy; Greece; Poland; Estonia; Lithuania; Moldova; Romania; Belarus; Ukraine; Russia: European Russia, Altai, Caucasus, Siberia, Amur, Primor’ye; Sakhalin Island; North Africa: Morocco, Algeria, Mayotte, Tunisia; Syria; Israel; Lebenon; Jordan; Turkey; Iraq; Iran; Afghanistan; Georgia; Azerbaijan; Armenia; Turmenistan; Uzbekistan; Tajikistan; Kazakhstan; Kyrgyzstan; Pakistan; India: Himachal Pradesh, Jammu and Kashmir; Nepal; Mongolia; Nei Menggu; China: Manchuria, Gansu, Hebei, Heilongjiang, Qinghai, Ningxia, Yunnan, Beijing, Liaoning, Xinjiang, Shaanxi; Korea. Introduced into Iceland; New Zealand; Australia (N. S. W., A. C. T., W. A., Vic., Tasmania); Ascension Island; South Africa; Chile; Argentina (Neuquén, Rio Negro, Chubut); Canada (Man., Ont., Que.); U. S. A. (NJ, NY, VT, MA, CT, DE, MD, PA, OH, IN, MI, WI, SD, VA, NC, WA, OR, CA, AZ).

inexspectata Eck
Vespula inexspectata Eck, 1994, Entomol. Abhand. Staat. Mus. Tierk. Dresden 56 (6): 125, female, figs. 1-6, 7 - “W. slope, Popocatepetl, Mx. MEX, 10000’“ (Ithaca); also from two other localities in México, and Michoacan.
Distribution: Mexico: México, Michoacan, Chiappas.

maculifrons (du Buysson)
Vespa maculifrons “S.” Harris, 1853, in Hitchcock, Rept. Geol. Mineral. Bot. Mass.: 589. Nomen nudum.
Vespa communis de Saussure, 1857, Stettin. Entomol. Ztg. 18: 116 (key), 117, female - “America septentr.” (Genève, Paris). Junior primary homonym of Vespa communis von Schrank, 1785.
Vespa maculifrons “H.” du Buysson, 1905 (1904), Ann. Soc. Entomol. Fr. 73: 608, female - [U. S. A.] “Delaware, Wilmington” (London), as a synonym of Vespa communis de Saussure, 1857. Available under Article 11.6.1 of the International Code of Zoological Nomenclature.
Vespa communis var. flavida Sladen, 1918, Ottawa Nat. 32: 71, female - “Common in eastern Canada, More southern range.” (type lost?).
Distribution: Canada: Man., Ont., Que., N. B., Nova Scotia; U. S. A.: MT, WY, CO, NM, TX, LA, MS, AL, AR, OK, KS, NE, ND, SD, MN, IA, MO, WI, IL, IN, MI, OH, PA, NY, VT, NH, ME, MA, CT, KY, NJ, DE, MD, VA, WV, KY, TN, NC, SC, GA, AL, FL; Mexico: Tamaulipas, Hidalgo, Nuevo León.

pensylvanica (de Saussure)
Vespa pensylvanica de Saussure, 1857, Stettin. Entomol. Ztg. 18: 116, 117, female - “America septentr., in Canada, et in montibus mexicanis” (lectotype from New Mexico, in Genève).
Vespa occidentalis Cresson, 1874, Trans. Am. Entomol. Soc. 5: 100, female, “male” [worker] - [U. S. A.] “Nevada” (lectotype female Philadelphia); also from NM. Junior primary homonym of Vespa occidentalis Olivier, 1792.
Distribution: Canada: B. C., Alta., Sask., Man.; U. S. A.: WA, OR, CA, NV, ID, MT, ND, SD, NE, MN, WI, WY, CO, UT, AZ, NM, TX; Mexico: Baja California Norte, México, Michoacan. Introduced into U. S. A. (HI); Australia: (Melbourne).

shidai Ishikawa, Yamane and Wagner
Vespula shidai Ishikawa, Yamane and Wagner, 1980, Ins. Matsum. 19: 1, 44, fig. 135A, female, male - “Sapporo, Hokkaidô, Japan” (holotype male Entomological Institute, Hokkaido University); also from other localities in Japan; and Russia.
Distribution: China: Heilongjiang, Liaoning, Manchuria; Korea; Russia: Amur, Primor’ye, Kurile Islands: Kunashir; Japan: Kyushu, Including Yakushima Island, Shikoku, Honshu, Hokkaido, Ryukyu Islands: Amami Island.

shidai amamiana Yamane
Vespula shidai amamiana Yamane, 1987, Kontyû 55 (4): 634 (key), 635, female, male - [Japan] “Naze, Amami-ôshima” (type depository unknown); also from elsewhere on Amami, and Kakeroma.
Distribution: Japan: Ryukyu Islands: Amami Island.

structor (Smith)
Vespa structor Smith, 1870, in Horne and Smith, Trans. Zool. Soc. Lond. 7 (3): 191, pl. XXI fig. 12, female - “Binsur, Kumaon, North-west Provinces of India” (London).
Vespa orbata var. aurulenta du Buysson, 1905 (1904), Ann. Soc. Entomol. Fr. 73 (3): 579, female - “Chine, Mon-pin; Amur” (Paris, Wien?).
Vespa structrix Schulz, 1906, Spolia Hym.: 231. Unjustified emendation of Vespa structor Smith.
Vespula gongshanensis Dong 2005, in Dong et al., Entotaxonomia 27 (1): 65, female, male – “China, Gongshan, Yunnan Province, alt. 2,950” (Holotype female (Yunnan).
Distribution: India: Jammu and Kashmir, Himachal Pradesh, Uttarakhand, Uttar Pradesh,West Bengal, Sikkim, Assam; Nepal; China: Gansu, Sichuan, Fujian, Shaanxi, Yunnan, Henan, Ningxia, Tibet; Myanmar; Thailand; Laos.

vulgaris (Linnaeus)
Vespa vulgaris Linnaeus, 1758, Syst. Nat., ed. 10, 1: 572 - “in Europa” (lectotype female, Linnaean Society, London).
Vespa sexcincta Panzer, 1799, Fauna Ins. German. 63: 1, fig. 1, male - “Viennae Austr.” (destroyed).
Vespa vulgaris var. pseudogermanica Stolfa, 1932, Mem. Soc. Entomol. Ital. 11 (1): 26 (key), female - [Italy] “Lago superiore di Weissenfels (Tarvisio) Senosecchia” (type depository unknown).
Distribution: Iceland; Norway; Sweden; Finland; Denmark; Germany; Netherlands; Belgium; Luxembourg; France, including Corsica; U. K.: England including Orkney Island, Isle of Man; Wales, Scotland including Shetland; Ireland; Spain; Portugal; Italy; Switzerland; Austria; Liechtenstein; Czech Republic; Hungary; Bosnia and Herzegovina; Slovenia; Albania; Romania; Macedonia; Greece; Croatia; Serbia; Bulgaria; Slovakia; Moldova; Poland; Belarus; Ukraine; Lithuania; Latvia; Estonia; Russia: European Russia to the Kola Peninsula, Altai, Caucasus, Siberia, Amur, Primor’ye; Azerbaijan; Georgia; Syria; Isreal; Turkey; Iran; Kazakhstan; Kyrgyzstan. China: Manchuria; Introduced into Iceland; New Zealand; Australia: (Victoria, Tasmania); Chile, Including Navarino Island; Argentina; South Africa: (Cape Town, St Helena); French Polynesia: (Tahiti Island).

vulgaris rufosignata Eck
Vespula rufosignata Eck, 1998, Reichenbachia 32 (43): 299, female, figs. 1A, C, 2 - “India: Kashmir, Gulmarg, 2900 m” (type depository not stated; London); also from two other localities.
Distribution: India: Jammu and Kashmir; Pakistan.

vulgaris vetus Eck
Vespula vulgaris vetus Eck, 1999, Entomol. Abhand. Staat. Mus. Tierk. Dresden 58: 309, 329, female, male - “China, Quinghai Prov., Datong-Tal, 120 km NE Xinging, 2360 m” (holotype female Dresden); also from five other localities.
Vespula yunlongensis Dong and Wang, 2003, in Dong et al., J. Southwest Agric. Univ. 25(3): 212, fig. 1, female [?] - “Yunnan, China” (Yunnan).
Distribution: Mongolia; Nei Menggu; China: Qinghai, Gansu, Sichuan, Shaanxi, Ningzia, Yunnan, Xinjiang, Heilongjiang, Beijing, Hebei, Liaoning.

vulgaris spp #A
Distribution: Japan: Hokkaido, Honshu. Russia: Sakhalin Island; Kurile Islands: Iturup, Kunashir.

Subgenus Rugovespula
Rugovespula Archer, 1982, Kontyû 50 (2): 261, 264, subgenus of Vespula Thomson.Type species: Vespa koreensis Radoszkowski, 1887, by original designation.

arisana (Sonan)
Vespa arisana Sonan, 1929, Trans. Nat. Hist. Soc. Formosa. 19 (101): 137 (key), 147, female - [Taiwan] “Fuukiko” (Taichung); also from elsewhere on Taiwan.
Distribution: Taiwan.

koreensis (Radoszkowski)
Vespa koreensis Radoszkowski, 1887, Horae Soc. Entomol. Ross. 21: 432, male, female - “Korée” (lectotype female London).
Pseudovespa birulai Gussakovskii, 1932, Ark. Zool. 24A (10): 52, female - [Russia] “Ussuri-Geb.: Tigrovaja; Sutshan, Sedanka, Kreis Chabarowsk, Wladiwostok” (Stockholm).
Distribution: N. China; Korea; Russia: Siberia, Amur, Primor’ye.

koreensis stizoides (Buysson)
Vespa germanica var. stizoides du Buysson, 1905 (1904), Ann. Soc. Entomol. Fr. 73: 494 (key), 615, female - “Nord de l’Inde” (London).
Distribution: India: Sikkim, Assam, Meghalaya; Laos; China: Yunnan.

koreensis salebrosa Archer
Vespula koreensis salebrosa Archer, 1982, Kontyû 50 (2): 261, 267, female - “China: Szechuen, Suifu” (London); also from numerous other localities in China.
Vespula hainanensis Lee, 1986, Sinozool. 10 (4): 201, 202, 206, fig. 1, female - [China] “Hainan Island, Guangdong Province” (Beijing).
Distribution: Myanmar; Thailand; Laos; Vietnam; China: Anhui, Sichuan, Yunnan, Heilongjiang, Beijing, Hubei, Hunan, Henan, Jiangxi, Liaoning, Zhejiang, Fujian, Shaanxi, Guangdong, Hong Kong, Hainan Island; Taiwan.

koreensis nujiangensis Dong & Wang
Vespula nujiangensis Dong and Wang, 2004, in Dong et al., J. Southwest Agric. Univ. (Nat. Sci.) 26 (2): 146, female, figs. 1-5 - [China] “Nujiang Liuku, Yunnan, Alt. 1,100m. 25°58’ N. 98°65’ E.” (Yunnan).
Distribution: China: Yunnan.

orbata (du Buysson)
Vespa orbata du Buysson, 1902, Bull. Soc. Entomol. Fr. 1902: 140, female - [India] “Dardjiling” (lectotype Paris); also from Sikkim; China.
Vespa minuta Dover, 1925 (1924), J. Asiat. Soc. Bengal (n. s.) 20: 304, fig. 9, female - [Myanmar] “Tenasserim” (London). Junior primary homonym of Vespa minuta Linnaeus, 1767.
Distribution: India: Assam, Uttarakhand, Uttar Pradesh, West Bengal; Nepal; Myanmar; Laos; Thailand; Vietnam; China: Tibet, Zhejiang, Hubei, Yunnan, Hainan Island, Fujian, Sichuan, Hunan, Heibei.
Anthony Daglio
 
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