A list of works on the social wasps of the Russian Far East

[Anthony Daglio]

I am compiling a list of the studies on the social wasps of the Russian Far east in this post in order to have a further understanding of these wasps on the whole. There are many interesting forms that exist in this area and hopefully this list will promote further interest in these wasps. In my opinion there are several species in the Genus Dolichovespula (D. pacifica Birula, D. pacifica loekenae Eck, D. norwegica peruana Saussure, D. saxonica kamtschatkensis Eck, and D. saxonica nigrescens Eck) Also three forms described by Birula (D. media lugubris Birula, D. adulterina colchica, and D. saxonica monticola) are of interest. I am not 100% if they are synonyms of other species. There are also there are a few forms in the Vespula rufa group that need to be reinvestigated (V. rufa spp# A (Undescribed form), Vespula schrenkii Radoszkowsi, and Vespula schrenkii spp#B (another undescribed form) which warrant special attention. I am adding images of these species.

#A GENERAL STUDIES ON THE SOCIAL WASPS OF THE RUSSIAN FAR EAST
1.) Sugihara (1939) A list of the speces of Vespinae from the Kurile Islands: https://www.academia.edu/1464981/Sugiha ... _Vespidae_
2.) Kurzenko (1995) A key to the species of Vespidae from the far east of Russia in: A key to insects of the Russian Far East Vol. 4. Part 1.: http://www.academia.edu/19513356/Kurzen ... _of_Russia
3.) Kurzenko (1995) Data to the fauna of the Aculeata of the Ussuri Area: http://www.academia.edu/11378835/Kurzen ... _Vespidae_
4.) Dubatolov (1998) Social wasps of Siberia in the collection of the Siberian Zoological Museum: http://www.academia.edu/1463783/Dubatol ... cal_Museum
5.) Dubatolov (1999) Social wasps of the Daurskii State Nature Reserve (No links to full text currently)
6.) Dubatolov et al (2002) New data on distribution of social wasps in Asian Russia and Northern China: http://www.academia.edu/1463821/Dubatol ... hern_China
7.) Kurzenko (2004) Vespid and Sapygid fauna of Sakhalin Island (No fulltext available at the moment).
8.) Dubatolov & Novomodnyi (2005) New data on distribution of social wasps in the Russian Far East: http://www.academia.edu/1463747/Dubatol ... n_far_east
9.) Dubatolov & Dolgick (2009) Social wasps of the Bolshekhekhtsirsky Nature Reserve (The Khabarovsk subburbs) with notes on their distribution in Lower Amur: http://www.academia.edu/1463764/Dubatol ... Lower_Amur
10.) Dubatolov (2011) Social wasps of the Bastak Nature Reserve (Jewish Autonomous Province) Amur Basin: http://www.academia.edu/1463755/Dubatol ... mur_Basin_
11.) Antropov & Khrustaleva (2009a) Species compositions, biological characteristics, and distribution of potentially dangerous wasps 1 (Vespinae: Vespa; Polistnae: Polistes): http://vespabellicosus2008.narod.ru/pdf7.pdf
12.) Antropov & Khrustaleva (2009b) Species compositions, biological characteristics, and distribution of potentially dangerous wasps 2 (Vespinae: Dolichovespula, Vespula): http://www.academia.edu/4301382/Antropo ... IN_RUSSIAN
13.) Smetanen (2013) On the insect fauna of the Kichiga river basin Northeasten Kamchatka: https://www.academia.edu/8886263/Smetan ... _Kamchatka
14.) Antropov & Prochalykin (2014) To the fauna of social wasps of Southern Siberia: http://www.academia.edu/7533856/Antropo ... rn_Siberia
15.) Abashev & Dabaev (2014) To the vespid fauna of National Park Alkhanai South-Eastern Transbalkalia: https://www.academia.edu/12564221/Abash ... sbaikalia_
16.) Dubatolov (2015) Social wasps of Asian Russia: http://www.academia.edu/3420429/Dubatol ... ian_Russia

#B WEBSITES ON THE SOCIAL WASPS OF ASIAN RUSSIA:
1.) Common wasps of Asian Russia: http://szmn.eco.nsc.ru/Vespidae/wasplist.htm
2.) Social wasps of Siberia in the collections of the SZM: http://szmn.eco.nsc.ru/vvdubat/pdf/ves_p1.htm
3.) Mobiol.ru: http://molbiol.ru/forums/index.php?act= ... chovespula
4.) Social wasps of Chuvahia and other parts of Russia: http://vespabellicosus2008.narod.ru/

#C CHECKLISTS OF THE SOCIAL WASPS OF THE RUSSIAN FAR EAST
1.) Kurzenko (2012) A checklist of the Vespidae of the Russian Far East in: Annotated catalog of the insects of the Russian Far East Vol.1 Hymenoptera: https://www.academia.edu/19513416/Kurze ... olistinae_
2.) Dubatolov (2015) Social wasps of Asian-Russia: http://www.academia.edu/3420429/Dubatol ... ian_Russia

#D TAXONOMIC AND FAUNISTIC REPORTS OF THE VESPINAE (VESPA, DOLICHOVESPULA, VESPULA) OF THE RUSSIAN FAR EAST
VESPA
1.) Birula (1925a) Sur la distribution geographique du frelon commun (Vespa crabro L.) et sur les races climatiques (No links to full text at the moment).
2.) Birula (1925b) Uber die Russischen wespen und ihre geographische verbreitung (No links to full text at the moment)
3.) Dubatolov et al (2003) Subspecies of Vespa crabro in two different papers by Birula 1925: http://www.academia.edu/1163340/Subspec ... la_in_1925
4.) Archer (2008) Taxonomy, Distribution and nesting biology of the Genera Provespa Ashmead and Vespa Linneaus: http://www.academia.edu/1468302/Archer_ ... _Vespidae_

DOLICHOVESPULA
1.) Birula (1927) Uber die Russischen wespen und ihre geogrphische verbreitung (Zweiter Bietrag): https://www.academia.edu/2455410/Birula ... r_Bietrag_
2.) Birula (1930) Uber die Russischen wespen und ihre geographische verbreitung (Dritter Beitrag):; http://www.academia.edu/2303103/Birula_ ... r_Beitrag_
3.) Yamane (1975) Taxonomic notes on the subgenus Boreovespula from Japan with notes on specimens from Sakhalin: http://www.academia.edu/1463575/Yamane_ ... m_Sakhalin
4.) Wagner (1978) On the Genus Dolichovespula and an addition to its known species in North America: http://www.academia.edu/1463501/Wagner_ ... _Vespidae_
5.) Eck (1981) Zur verbreitung und variabilitat von Dolichvespula norwegica: http://www.academia.edu/1464834/Eck_198 ... _IN_GERMAN
6.) Eck (1983) Zur verbreitung und variabilitat von Dolichovespula saxonica: http://www.academia.edu/1463505/Eck_198 ... _IN_GERMAN
7.) Eck (1984a) Bestimmungschlussel fur die Arten der Gattung Dolichovespula Rowher: http://www.academia.edu/1464843/Eck_198 ... _IN_GERMAN
8.) Eck (1984b) Uber die verbreitung von Dolichovespula pacifica Birula und den status von Dolichovespula xanthicincta Archer: http://www.academia.edu/1464839/Eck_198 ... _IN_GERMAN
9.) Eck (1984c) Zur verbitung von Dolichovespula loekenae Eck und ihrer stellung zu den nachstverwanden Anten: http://www.academia.edu/1505810/Eck_198 ... _IN_GERMAN
10.) Pekarinen (1995) Geographic variation and taxonomy of the species of Dolichovespula in the boreal zone of the Holartic region: http://www.academia.edu/1464905/Pekarin ... _vespinae_
11.) Archer (1999) Taxonomy and distribution of the Euro-Asian species of the Geunus Dolichovespula: http://www.academia.edu/1463405/Archer_ ... _Vespidae_
12.) Archer (2006) Taxonomy, distribution and nesting biology of the species of the Genus Dolichovespula: http://www.academia.edu/1463442/Archer_ ... _Vespidae_
13.) Carpenter et al (2015) Taxonomic notes on Vespinae: http://www.academia.edu/16840334/Carpen ... _Vespidae_

VESPULA
1.) Birula (1930) Uber die Russischen und ihre geographische verbreitung (Vierter Breitrag): http://www.academia.edu/2303096/Birula_ ... _Breitrag_
2.) ) Yamane et al (1980) A tentative revision of the subgenus Paravespula of eastern Asia: http://www.academia.edu/1463552/Yamane_ ... _Vespidae_
3.) Archer (1981) The Euro Asian species of the Vespula rufa group with descriptions of two new species and one subspecies: http://www.academia.edu/1463618/Archer_ ... subspecies
4.) Archer (1982) A revision of the subgenus Rugovespula of the Genus Vespula: http://www.academia.edu/1463550/Archer_ ... _Vespidae_
5.) Archer (1997) Taxonomy distribution and nesting biology of the Euro-Asian species of the Vespula rufa group: http://www.academia.edu/1463411/Archer_ ... _Vespidae_
6.) Archer (1998) The world distribution of the species Euro-Asian of the Genus Paravespula: http://www.academia.edu/1463407/Archer_ ... _Vespidae_
7.) Archer (2007) Taxonomy distribution and nesting biology of the Genus Vespula or the Vespula rufa group: http://www.academia.edu/1463444/Archer_ ... _Vespidae_
8.) Archer (2008) Taxonomy distribution and nesting biology of the Genus Paravespula or the Vespula vulgaris group: http://www.academia.edu/1463476/Archer_ ... _Vespidae_

#E CHECKLIST OF THE VESPINAE SPECIES OF THE RUSSIAN FAR EAST
VESPA
a.) Vespa analis Fabricius: http://szmn.eco.nsc.ru/Vespidae/analis.htm
b.) Vespa binghami Buysson: http://szmn.eco.nsc.ru/Vespidae/binghami.htm
c.) Vespa crabro Linnaeus: http://szmn.eco.nsc.ru/Vespidae/crabro.htm
d.) Vespa ducalis Smith: http://szmn.eco.nsc.ru/Vespidae/ducalis.htm
e.) Vespa dybowskii Smith: http://szmn.eco.nsc.ru/Vespidae/dybowskii.htm
f.) Vespa mandarinia Smith: http://szmn.eco.nsc.ru/Vespidae/mandarinia.htm
g.) Vespa simillima Smith: http://szmn.eco.nsc.ru/Vespidae/simillima.htm

DOLICHOVESPULA
a.) Dolichovespula adulterina Buysson: http://szmn.eco.nsc.ru/Vespidae/adulterina.htm
b.) Dolichovespula media Retzius: http://szmn.eco.nsc.ru/Vespidae/media.htm
c.) Dolichovespula norwegica Fabricius: http://szmn.eco.nsc.ru/Vespidae/norwegica.htm
d.) Dolichovespula norwegica peruana Saussure: http://www.academia.edu/4986473/Dolicho ... sure_1867_
e.) Dolichovespula omissa Bischoff: http://szmn.eco.nsc.ru/Vespidae/omissa.htm
f.) Dolichovespula pacifica Birula: http://szmn.eco.nsc.ru/Vespidae/pacifica.htm http://www.academia.edu/4987234/Dolicho ... irula_1930
g.) Dolichovespula pacifica loekenae Eck: http://www.academia.edu/4987404/Dolicho ... e_Eck_1980
h.) Dolichovespula saxonica Fabricius: http://szmn.eco.nsc.ru/Vespidae/saxonica.htm http://www.academia.edu/4987607/Dolicho ... cius_1793_
i.) Dolichovespula saxonica kamtchatkensis Eck: http://www.academia.edu/4986687/Dolicho ... s_Eck_1983
j.) Dolichovespula saxonica nigrescens Eck: http://www.academia.edu/4986653/Dolicho ... s_Eck_1983
k.) Dolichovespula sylvestris Scopoli: http://szmn.eco.nsc.ru/Vespidae/sylvestris.htm

VESPULA
a.) Vespula austriaca Panzer: http://szmn.eco.nsc.ru/Vespidae/austriaca.htm
b.) Vespula flaviceps Smith: http://szmn.eco.nsc.ru/Vespidae/flaviceps.htm
c.) Vespula germanica Fabricius: http://szmn.eco.nsc.ru/Vespidae/germanica.htm
d.) Vespula flaviceps Smith: http://szmn.eco.nsc.ru/Vespidae/flaviceps.htm
e.) Vespula koreensis Radoszkowski: http://szmn.eco.nsc.ru/Vespidae/koreensis.htm
f.) Vespula rufa Linnaeus: http://szmn.eco.nsc.ru/Vespidae/rufa.htm http://www.academia.edu/4996697/Alloves ... aeus_1758_
g.) Vespula rufa Spp#A: http://www.academia.edu/4996726/Allovespula_rufa_Spp_A
h.) Vespula schrenkii Radoszkowski: http://www.academia.edu/4996801/Alloves ... wski_1861_
i.) Vespula schrenkii siberica Andre: http://www.academia.edu/4996812/Alloves ... ndre_1884_
j.) Vespula schrenkii Spp#B: http://www.academia.edu/4996841/Alloves ... nkii_Spp_B
k.) Vespula shidai Ishikawa, Yamane, & Wagner: http://szmn.eco.nsc.ru/Vespidae/shidai.htm
l.) Vespula vulgaris Linnaeus: http://szmn.eco.nsc.ru/Vespidae/vulgaris.htm

#F TAXONOMIC & FAUNISTIC STUDIES ON POLISTES SPECIES OF THE RUSSIAN FAR EAST
1.) Yamane Sk. & Yamane So. (1987) A new species and new synonomy in the subgenus Polistes of eastern Asia: http://www.researchgate.net/publication ... stern_Asia
2.) Dubatolov (1998) Social wasps of Siberia in the collection of the SZM: http://www.academia.edu/1463783/Dubatol ... cal_Museum
3.) Social wasps of the Daurskii State Nature Reserve (Southeast Siberia) (No links to fulltext available)
4.) Dubatolov et al (2002) New data on distribution of social wasps in Asian Russia and northern China: http://www.academia.edu/1463821/Dubatol ... hern_China
5.) Dubatolov & Novomodnyi (2005) New data on distribution of social wasps in the Russian far east: http://www.academia.edu/1463747/Dubatol ... n_far_east
6.) Dubatolov & Dolgick (2009) Social wasps of the Bolshekhekhtsirski Nature Reserve (The Khabarovsk Suburbs) with notes on thier distribution in Lower Amur: http://www.academia.edu/1463764/Dubatol ... Lower_Amur
7.) Dubatolov (2011) Social wasps of the Bastak Nature Reserve (Jewish Autonomous Province) Amur Basin: http://www.academia.edu/1463755/Dubatol ... mur_Basin_
8.) Dubatolov et al (2015) The Rothneyi's wasp, Polistes rothneyi Cameron 1900, A new species for the Russian fauna from Southern Primor'ye: https://www.researchgate.net/publicatio ... urnal_T_VI
#G CHECKLIST OF THE POLISTES SPECIES OF THE RUSSIAN FAR EAST
POLISTES
a.) Polistes biglumis Linnaeus: http://szmn.eco.nsc.ru/Vespidae/biglumis.htm
b.) Polistes chinensis Fabricius: http://szmn.eco.nsc.ru/Vespidae/chinensis.htm
c.) Polistes dominula Christ: http://szmn.eco.nsc.ru/Vespidae/dominulus.htm
d.) Polistes gallicus Linnaeus: http://szmn.eco.nsc.ru/Vespidae/gallicus.htm
e.) Polistes nimpha Christ: http://szmn.eco.nsc.ru/Vespidae/nimpha.htm
f.) Polistes riparius Yamane & Yamane: http://szmn.eco.nsc.ru/Vespidae/riparius.htm

POLISTELLA
a.) Polistes diakonovi Kostylev: http://szmn.eco.nsc.ru/Vespidae/diakonovi.htm
b.) Polistes senellini Saussure: http://szmn.eco.nsc.ru/Vespidae/snelleni.htm

GYROSTOMA
a.) Polistes rothneyi Cameron

[Anthony Daglio]

Here are images of Dolichovespula pacifica Birula from the Russian far east and Japan.

[Anthony Daglio]

Here are some images of Dolichovespula pacifica loekenae Eck. Matsuura & Yamane (1990) and Pekkarinen (1995) considered D. pacifica and D. loekenae to be subspecies of Dolichovespula norvegicoides and Archer (1989, 1999, 2006, 2012) consider loekenae a subspecies of pacifica even though Eck (1984) and Pekkarinen found differences in morphology and male genetalia. Recently Persson (2015) found genetic differences between populations of D. pacifica and D. loekenae. I consider the two to be distinct forms. D. loeknae is stable in color pattern throughout its entire range which is quite widespread and so is D. pacifica which is restricted in its distribution. Plus both forms do not overlap in distribution and are disjunct populations. I think both of these forms warrant reinvestigation.

[Anthony Daglio]

Here are some photos of the nominate form of Dolichovespula saxonica Fabricius to compare with the subspecies. The nominate form is found in European Russia but is replaced by Dolichovespula saxonica nigrescens Eck in the Russian far east and Dolichovespula saxonica kamtchatkensis Eck in Kamchatka.

[Anthony Daglio]

Here is the subspecies Dolichovespula kamtchatkensis which has a very restricted distribution in Kamchatka and Commander Island in the Russian Far East.

[Anthony Daglio]

Here are a few images of Dolichovespula saxonica nigrescens which is a common wasp in the Russian far east.

[Anthony Daglio]

Here is the nominate species of vespula rufa Linnaeus which is widespread in Europe and European Russia.

[Anthony Daglio]

Here is an undescribed species or subspecies that is found in Norway, Sweden, Finland, Hungary across to the Russian Far east. It differs from the nominate form of rufa in having the markings of the head, thorax, and first abdominal tergite being ivory white and the rest of the abdominal terga are dull yellow. Archer (1997) briefly described workers of this form from Sweden.

[Anthony Daglio]

Here are some photos of Vespula schrenkii Radoszkowski, A species found in Korea, China, Mongolia, and the Russian Far East. In the coarse of my studies I have decided to treat this form as a distinct species. In Eastern Asia it has several subspecies or species closely related to it (Daglio Unpublished).

[Anthony Daglio]

Here are some images of Vespula schrenkii siberica Andre. This form is found in Siberia? Japan, Kunishir, Sakhalin, and the Kurile Islands.

[Anthony Daglio]

Here is an undescribed from of Vespula schrenkii that is found in Afghanistan, Uzbekistan, Tajikistan, Kazakhstan, Kyrgyhzstan, Mongolia, China, Japan (rarely), The Russian Far East.

[Anthony Daglio]

As can clearly be seen a taxonomic revision of some forms of Vespula and Dolichovespula in the Russian far east is warranted.